I downgraded my nüvi 255W from FW 7.60 to 5.00 without any problem.
Reason I needed FW 5.00 is that I wanted to establish in which FW version the support for map files larger than 2 GB was added. I can now confirm that this happened as of FW 5.10 dated 8 June 2009 (FW 5.00 does not yet support maps > 2 GB).
This support not mentioned anywhere in the change log. I wonder why, since the change log of many other models does mention it, for instance nüvi 5xx with FW 3.40, or nüvi 8xx with FW 4.70, or zumo 4xx / 5xx with FW 4.90 plus a collection of older handheld outdoor models. An important enhancement like the support of file sizes > 2 GB belongs in the change log, also of the nüvi 2x5 and 2x5W...
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